[PR #1837/4cf073bb backport][0.20] Fix import collection marks and signatures #1840
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This is a backport of PR #1837 as merged into main (4cf073b).
The import code was unable to link up a collection version with its metadata because using multiple fields as a foreign key is not supported by django-import-export. The only viable fallback given existing export files is to use the upstream_id.
fixes #1836